Parnara

Parnara Moore, [1881]

Parnara Moore, [1881] (Hesperiidae); Lepid. Ceylon 1 (4): 166; TS: Eudamus guttatus Bremer & Grey
Baorynnis Waterhouse, 1932; Aust. Zool. 7 (3): 201; TS: Pamphila amalia Semper
- = Parnara; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 179

Baorynnis; [NHM-BMW]
Parnara; [BMP]; [LepIndex]; [CoL] ; [iNaturalist] ; [NHM-BMW]
Parnara (Hesperiinae); [BK], 435; [AfrL]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 75; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 13; [RFEL]

Parnara guttatus (Bremer & Grey, [1852])

Ceylon, India, Burma, N.China - Himalayas, Afghanistan - Pakistan, Japan, Indonesia. See [maps] Straight Swift

Eudamus guttatus Bremer & Grey, [1852]; in Motschulsky, Etud. Ent. 1: 60; TL: Beijing, China
Eudamus guttatus Bremer & Grey, 1853; Schmett. N. China: 10, pl. 3, f. 2

Parnara guttata; [MRS], 727; Kudrna, 1974, Atalanta 5: 118; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181, f. 13; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 13; [iNaturalist]
Parnara guttatus; [BRU], 132; [BIR], 486 (key); [BOW]: pl. 188, f. 6; [RFEL]; [LepIndex] ; [CoL] ; [LEPIFORUM] ; [Yutaka Inayoshi]

800x1041 (~90Kb) upperside Japan: Lake Kawaguchiko, Yamanashi Prefecture, 23.8.2007, Photo © Peter Payzant

640x832 (~115Kb) underside Japan: Lake Kawaguchiko, Yamanashi Prefecture, 23.8.2007, Photo © Peter Payzant

700x1058 (~88Kb) upperside Japan: Tokyo, 1.9.2007, Photo © Peter Payzant

P. g. guttatus N.China - Japan

Hesperia fortunei Felder, 1862; Verh. zool.-bot. Ges. Wien 12 (1/2): 489; TL: Shanghai
- = Parnara guttata guttata; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181
Hesperia dalima Plötz, 1883; Stett. ent. Ztg 44 (1-3): 44; TL: "Brazilien"
- = Parnara guttata guttata; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181
Parnara kotoshona Sonan, 1936; Zephyrus 6 (3/4): 214; TL: Formosa
- = Parnara guttata guttata; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181
- = Parnara guttata; Chiba, Hsu, Tsukiyama, Lo, Chen & Wang, 2009, Zootaxa 2202: 53

Parnara guttata guttata; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181; [iNaturalist]

P. g. mangala (Moore, [1865]) S.China, W.China - Himalayas, Afghanistan, Pakistan

Pamphila mangala Moore, [1865]; 792; TL: N.India
Pamphila philino Möschler, 1878; 220; TL: Formosa
- = Parnara guttata mangala; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181
Hesperia kolantus Plötz, 1885; Berl. ent. Z. 29 (2): 227; TL: Inida
- = Parnara guttata mangala; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181
Pamphila ormuzd Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4): 307; TL: Kabadian
- = Parnara guttata mangala; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181

Pamphila ormuzd; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 507, pl. 18, f. 5
Parnara guttata mangala; [MRS], 727; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181
Parnara guttata ormuzd;
Parnara guttatus mangala; [iNaturalist]
Parnara philino; Piepers & Snellen, 1910, Rhop. Java [2]: 34, pl. 8, f. 48a-b
?Carterocephalus ormuzd; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 12

Parnara batta Evans, 1949

SE.China (Fukien, ...). See [maps]

Parnara guttatus batta Evans, 1949; Cat. Hesp. Europe Asia Australia Brit. Mus.: 433; TL: Fukien
- = Parnara guttata guttata; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 181

Parnara batta; Devyatkin & Monastyrskii, 2002, Atalanta 33 (1/2): 150, pl. Va, f. 9-10; [Yutaka Inayoshi]
Parnara guttata batta; [iNaturalist]
Parnara guttatus batta; [LepIndex]; [CoL]

Parnara ogasawarensis Matsumura, 1906 Bonin Is.

Parnara ogasawarensis Matsumura, 1906; Annot. zool. japon. 6 (1): 13, pl. 1, f. 4; TL: Bonin Is.

Parnara guttata ogasawarensis; Evans, 1949
Parnara guttatus ogasawarensis; [LepIndex]; [CoL]
Parnara ogasawarensis; Shirôzu, 1953; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 182, f. 14; [iNaturalist]

Parnara apostata (Snellen, [1880])

-. See [maps]

Pamphila apostata Snellen, [1880]; Midden-Sumatra, 4 (8): 27; TL: Borneo, Sarawak

Parnara apostata; Piepers & Snellen, 1910, Rhop. Java [2]: 41, pl. 9, f. 59; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 182; [iNaturalist]
Parnara guttata apostata; [iNaturalist]
Parnara guttatus apostata; [LepIndex]; [CoL]

P. a. apostata Sumatra, Malay Peninsula, S.Burma, Java, Bali

Hesperia haga Plötz, 1886; Stettin ent. Ztg 47 (1-3): 96; TL: Java
- = ; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 182
Hesperia sifa Plötz, 1886; Stettin ent. Ztg 47 (1-3): 96; TL: Java
- = ; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 182

Parnara apostata apostata; [BMP]: 385, pl. 60, f. 3-4; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 182, f. 1-3, 15-16; [iNaturalist] ; [Yutaka Inayoshi]

P. a. andra Evans, 1949 Borneo

Parnara guttatus andra Evans, 1949; Cat. Hesp. Europe Asia Australia Brit. Mus.: 434; TL: Borneo, Sarawak

Parnara apostata andra; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 182, f. 7, 17; [iNaturalist]
Parnara guttata andra; [iNaturalist]
Parnara guttatus andra; [LepIndex]; [CoL]

P. a. debdasi Chiba & Eliot, 1991 Nepal

Parnara apostata debdasi Chiba & Eliot, 1991; Tyô to Ga 42 (3): 183, f. 4-6, 18; TL: Nepal, Kaski District, Pothana

P. a. hulsei Devyatkin & Monastyrskii, 1999 S.Yunnan, ...

Parnara apostata hulsei; Huang & Xue, 2004, Neue Ent. Nachr. 57: 146 (name); [Yutaka Inayoshi]

Parnara kawazoei Chiba & Eliot, 1991

Philippines (Luzon), Borneo, Sulawesi. See [maps]

Parnara kawazoei Chiba & Eliot, 1991; Tyô to Ga 42 (3): 183, f. 8-11, 19, 20; TL: Philippines, Luzon

Parnara kawazoei; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 76; [iNaturalist]

Parnara ganga Evans, 1937

S.India - Vietnam - S.China, Malay Peninsula. See [maps] Continental Swift

Parnara ganga Evans, 1937; Entomologist 70 (4): 83; TL: Manipur

Parnara ganga; [MRS], 728; [BMP]: 386, pl. 60, f. 5; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186, f. 21; [LepIndex] ; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]

Parnara bada (Moore, 1878)

Vietnam. See [maps] Water Watchman Ceylon Swift

Hesperia bada Moore, 1878; Proc. zool. Soc. Lond. 1878 (3): 688; TL: Ceylon

Parnara bada; Moore, [1881], Lepid. Ceylon 1 (4): 167, pl. 70, f. 2, 2a; [BMP]: 385, pl. 60, f. 1-2; [MRS], 728; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186, f. 21; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 76; [iNaturalist] ; [LEPIFORUM] ; [Don Herbison-Evans]

602x729 (~57Kb) underside Thailand, Chon Buri Province, Pattaya, a small wood at a grassy swamp at the Jomtien beach, between B.O. Guesthouse and Jomtien Metro Condotel, 2nd January, 2006, Photo © Oleg Kosterin

P. b. bada Sri Lanka, India, China, Indo-China, Malay Peninsula, Sumatra, Java, Lesser Sunda Is., Borneo, Palawan, Philippines, Sulawesi

Hesperia quinigera Moore, 1878; Proc. zool. Soc. Lond. 1878 (3): 703; TL: Hainan
- = ; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186
Gegenes hainanus Moore, 1878; Proc. zool. Soc. Lond. 1878 (3): 703; TL: Hainan
- = ; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186
Hesperia intermedia Plötz, 1883; Stett. ent. Ztg 44 (1-3): 44; TL: Java
- = ; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186
Hesperia daendali Plötz, 1885; 226; TL: Batavia
- = ; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186
Hesperia nondoa Plötz, 1886; Stettin ent. Ztg 47 (1-3): 97; TL: Manila
- = ; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186
Baoris distictus Holland, 1887; Trans. Amer. Ent. Soc. 14: 123; TL: Hainan
- = ; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186
Baoris (Parnara) philotas de Nicéville, 1895; J. Bombay nat. Hist. Soc. 9 (4): 402, (3): pl. Q, f. 60; TL: North Kanara; Trevandrum, S.India
- = ; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186

Parnara bada bada; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 76; [Yutaka Inayoshi]
Parnara naso bada; Kudrna, 1974, Atalanta 5: 118; [LepIndex]; [CoL]

P. b. sida (Waterhouse, 1934) N.Queensland

Baoris sida Waterhouse, 1934; 415; TL: N.Queensland

Parnara bada sida; Chiba & Eliot, 1991, Tyô to Ga 42 (3): 186; [iNaturalist]
Parnara naso sida; ; [LepIndex]

P. b. borneana Chiba & Eliot, 1991 Borneo (Sabah)

Parnara bada borneana Chiba & Eliot, 1991; Tyô to Ga 42 (3): 187; TL: Sabah, Mt Marapok

Parnara bada borneana; [iNaturalist]

Parnara naso (Fabricius, 1798)

Africa. See [maps]

Hesperia naso Fabricius, 1798; Ent. Syst. (Suppl.): 431, no. 236-7
Hesperia marchalii Boisduval, 1833; Faun. ent. Madagascar, Lépid.: 66

Parnara marchalii; Evans, 1937, Cat. African Hesp. Brit. Mus.: -
Parnara naso podora; [LepIndex]; [CoL]
Parnara naso; Dunn & Dunn, 1991, Review Austr. Butts.:; [AfrL]; [LepIndex] ; [CoL] ; [iNaturalist]

P. n. naso

Parnara naso naso; [AfrL]; [iNaturalist]

P. n. poutieri (Boisduval, 1833) Madagascar

Hesperia poutieri Boisduval, 1833; Faun. ent. Madagascar, Lépid.: 65

Parnara naso poutieri; [MadL]; [AfrL]; [LepIndex]; [CoL] ; [iNaturalist]
Parnara poutieri; Evans, 1937, Cat. African Hesp. Brit. Mus.: -

P. n. bigutta Evans, 1937

Parnara naso bigutta; [AfrL]; [LepIndex]; [CoL] ; [iNaturalist]

Parnara monasi (Trimen, 1889)

Natal, Transvaal, Moçambique. See [maps] Water Watchman

Pamphila monasi Trimen, 1889; South-African Butts 3: 317
Pamphila neoba Mabille, 1891; Bull. Soc. Ent. Belg. 35 (18): clxxviii; TL: Cameroons
Parnara anelia Bethune-Baker, 1908; Ann. Mag. nat. hist. (8) 2 (12): 480; TL: Mawamba-Makala

Parnara monasi; Evans, 1937, Cat. African Hesp. Brit. Mus.: -; [AfrL]; [iNaturalist]
Parnara monazi[sic]; [BOW]: pl. 130, f. 9
Parnara naso monasi; [BK]: 436, pl. 64, f. 868; [LepIndex]; [CoL]

Parnara amalia (Semper, [1879])

Queensland. See [maps]

Pamphila amalia Semper, [1879]; J. Mus. Godeffroy 5 (14): 183, no. 154
Hesperilla fulgidus Miskin, 1889

Parnara amalia; Dunn & Dunn, 1991, Review Austr. Butts.:; [LepIndex] ; [CoL] ; [iNaturalist] ; [Don Herbison-Evans]
Parnara amalis[sic]; [BOW]: pl. 188, f. 5

Parnara kuwanoi Seok, 1937; Annot. zool. jap. 16 (2): 112; TL: Peiping

Hesperia wambo Plötz, 1886; Stettin ent. Ztg 47 (1-3): 97; TL: Africa

References:

[AfrL] Mark C. Williams
Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[CiNii]
http://ci.nii.ac.jp/
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[Don Herbison-Evans] and Stella A. Crossley
Australian Caterpillars and their Butterflies and Moths
[EVERSMANNIA]
Eversmannia contents
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[LepIndex] Global Lepidoptera Index
Beccaloni, G., Scoble, M., Kitching, I., Simonsen, T., Robinson, G., Pitkin, B., Hine, A., Lyal, C., Ollerenshaw, J., Wing, P., & Hobern, D. (2024). Global Lepidoptera Index. In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, L. Vandepitte, D. Hobern, P. Schalk, R. E. DeWalt, K. Ma, J. Miller, T. Orrell, R. Aalbu, J. Abbott, R. Adlard, C. Aedo, et al., Catalogue of Life Checklist (1.1.24.106); https://doi.org/10.48580/dg4lg-49xk
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Peter Payzant]
Canada; http://www.pbase.com/ppayzant/profile
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Bethune-Baker, 1908
Descriptions of new Rhopalocera from the Upper Congo Ann. Mag. nat. hist. (8) 2 (12) : 469-482
Boisduval, 1833
Faune Entomologique de Madagascar, Bourbon et Maurice. Lépidoptéres Faun. ent. Madagascar, Lépid. : 1-122, pl. 1-16
Bremer & Grey, [1852]
Diagnoses de Lépidopterères nouveaux, trouvés par MM. Tatarinoff et Gaschkewitch aux environs de Pekin in Motschulsky, Etud. Ent. 1 : 58-67
Bremer & Grey, 1853
Beiträge zur Schmetterlings-Fauna des nördlichen China Schmett. N. China : 5-23
Chiba & Eliot, 1991
A revision of the genus Parnara Moore (Lepidoptera, Hesperiidae) with special reference to the Asian species Tyô to Ga 42 (3) : 179-194
Chiba, Hsu, Tsukiyama, Lo, Chen & Wang, 2009
Jinhaku Sonan's skipper type collection deposited at Taiwan Agricultural Research Institute (Lepidoptera: Hesperiidae) Zootaxa 2202 : 48-58
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2 Mon. Rhop. Sinesium : 1-854
[BMP]; Corbet & Pendlebury, 1992
The butterflies of the Malay Peninsula. Fourth Edition revised by J. N. Eliot with plates by Bernard D'Abrera Butterflies Malay Peninsula (4th edn) : 1-595, pl. 1-69
de Nicéville, 1895
On new and little-known Butterflies from the Indo-Malayan Region J. Bombay nat. Hist. Soc. 9 (3): 259-321, pl. N-Q, (4): 366-410
Devyatkin & Monastyrskii, 2002
Hesperiidae of Vietnam 12: A further contribution to the Hesperiidae fauna of North and Central Vietnam Atalanta 33 (1/2) : 137-155
Dunn & Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy Review Austr. Butts. (1): 1-196, (2): 197-335, (3): 336-512, (4): 513-660
Evans, 1937
Indo-Australian Hesperiidae: Description of new Genera, Species and Subspecies Entomologist 70 (1): 16-19 (part 9), (2): 38-40 (part 10), (3): 64-66 (part 11), (4): 81-83 (part 12)
Evans, 1937
A catalogue of the African Hesperiidae. British Museum Cat. African Hesp. Brit. Mus. : 1-212, pl. 1-30
Evans, 1949
A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum Cat. Hesp. Europe Asia Australia Brit. Mus. : 1-502, pl. 1-53
Fabricius, 1798
Supplementum Entomologiae Systematicae Ent. Syst. (Suppl.) : 1-572, (index) 1-53
Felder, 1862
Verzeichniss der von den Naturforschern der k. k. Fregatte Novara gesammelten Macropepidoteren Verh. zool.-bot. Ges. Wien 12 (1/2) : 473-496
Grum-Grshimailo, 1888
Novae species et varietates Rhopalocerum e Pamir Horae Soc. ent. Ross. 22 (3-4) : 303-307
Grum-Grshimailo, 1890
Le Pamir et sa faune lépidoptérologique in Romanoff, Mém. Lép. 4 : 1-576, pl. A,1-21
Holland, 1887
Notes upon small collection of Rhopalocera made by Rev. B. C. Henry in the Island of Hainan, together with descriptions of some apparently new species Trans. Amer. Ent. Soc. 14 : 111-124
Huang & Xue, 2004
A Contribution to the Butterfly Fauna of Southern Yunnan Neue Ent. Nachr. 57: 135-154, pl. 11
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
Kudrna, 1974
An annotated List of Japanese Butterflies Atalanta 5 : 92-120
[BK]; Larsen, 1996
The Butterflies of Kenya and their natural history Butterflies of Kenya
[MadL]; Lees, Kremen & Raharitsimba, 2003
Classification, Diversity and Endemism of the Butterflies (Papilionoidea and Hesperioidea): A Revised Species Checklist in Goodman & Benstead, Nat. Hist. Madagascar : 762-793
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812
[BOW]; Lewis, 1974
Butterflies of the World
Mabille, 1891
Description d'Hespérides nouvelles (1-3) Bull. Soc. Ent. Belg. 35 (16): lix-lxxxviii, (17): cvi-cxxi, (18): clxviii-clxxxvii
Matsumura, 1906
Neue Rhopaloceren Japans Annot. zool. japon. 6 (1) : 7-15, pl. 1
Moore, 1878
Descriptions of new Asiatic Hesperidae Proc. zool. Soc. Lond. 1878 (3) : 686-695, pl. 45
Moore, 1878
List of Lepidopterous Insects collected by the late R. Swinhoe in the Island of Hainan Proc. zool. Soc. Lond. 1878 (3) : 695-708
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Piepers & Snellen, 1910
The Rhopalocera of Java. Hesperidae Rhop. Java [2] : 1-60, pl. 5-10
Plötz, 1883
Die Hesperiinen -- Gattung Hesperia Aut. und ihre Arten. (3) & (4) Stett. ent. Ztg 44 (1-3): 26-64, (4-6): 195-233
Plötz, 1885
Neue Hesperiden des indischen Archipels und Ost-Africa's aus der Collection des Herrn H. Ribbe in Blasewitz-Dresden, gesammelt von den Herren: C. Ribbe auf Celebes, Java un den Aru-Inseln, Künstler auf Malacca (Perak); Kühn auf West-Guinea (Jekar); Menger auf Ceylon Berl. ent. Z. 29 (2) : 225-232
Plötz, 1886
Nachtrag und Berichtigungen zu den Hesperiinen Stettin ent. Ztg 47 (1-3) : 83-117
Semper, [1879]
Beitrag zur Rhopalocerenfauna von Australien J. Mus. Godeffroy 5 (14) : 138-194, pl. 8-9
Seok, 1937
Prof. H. Kuwano's collection of butterflies from China Annot. zool. jap. 16 (2) : 106-115
Snellen, [1880]
Natuurlijke Historie. Achtse afdeeling. Lepidoptera door P. C. T. Snellen, met eene inleiding door Joh. F. Snelleman. Midden-Sumatra Midden-Sumatra, 4 (8) : 1-84, pl. 1-2 (?1880, ?1884, [1886]), : 85-92, pl. 3-5 ([1892])
Sonan, 1936
Notes on some butterflies from Formosa. (4) Zephyrus 6 (3/4) : 205-216
Trimen, 1889
South-African Butterflies: a monograph of the extra-tropical species. Vol. 3 South-African Butts 3 : 1-438, pl. 10-12
[BRU]; Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997
Guide to the Butterflies of Russia and adjacent territories: Hesperiidae, Papilionidae, Pieridae, Satyridae Butts. Russia adj. terr. 1
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Waterhouse, 1932
New genera of Australian Hesperiidae and a new subspecies Aust. Zool. 7 (3) : 198-201
[BIR]; Wynter-Blyth, 1957
Butterflies of the Indian Region (1982 Reprint)

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